Trifurcula immundella

Diagnostic description: 

Diagnosis.  T. immundella is most likely to be confused with the  slightly smaller T. serotinella;  small specimens found in localities where  the hosts of both species occur can only be identified with certainty by  their genitalia.  Other similar species, T. beirnei and T. squamatella  can be identified by their larger size and number of antennal segments.  The smaller T. ortneri is more ochreous and has a hair-pencil on the  hindwing (in male).  The male genitalia differ from T. serotinella by  the wider capsule and valva, the presence of anterior processes of the  gnathos and longer sublateral processes.

Morphology: 

Male. Wingspan:  6.0-8.2 mm.  Head: frontal tuft varying  from yellowish white to mixed ferruginous fuscous, sometimes almost  completely white;  collar paler, yellowish white;  scape white, usually  with few to several brown scales;  antenna with 35-43 segments.  Forewing  pale ochreous grey to greyish brown, irrorated by presence of white and  dark tipped scales;  colour very variable, extremes from almost completely  white to almost completely brown occur;  cilia-line irregular, terminal  cilia white.  Hindwing: greyish white.  Abdomen with yellowish anal tufts. Female.  Wingspan:  6.4-8.4 mm.  Antenna with 34-40 segments.
Male genitalia. Vinculum anteriorly truncate.  Uncus  truncate, not widened.  Gnathos with broadly triangular central element,  ending in a rounded tip;  with narrow diverging anterior processes.  Valva with margins at base parallel, beyond middle margins suddenly narrow  towards slightly curved tip;  dorsally excavated over more than half its  length at base;  transtilla with straight transverse bar.  Aedeagus  slender, tip bifurcate;  with one large straight, spine-like cornutus and  two large curved cornuti with large base. Female genitalia. T VIII covered with many scales and several  setae;  anal papillae each with 18-23 setae;  apophyses curved.  Vestibulum  small, folded.  Ductus bursae with folds, long;  corpus bursae elongate,  covered with minute spines and pectinations;  signa long and narrow,  margins irregular, often with incomplete cells.  Ductus spermathecae with  4½ convolutions.

Associations: 

Host plants: Cytisus scoparius (= Sarothamnus scoparius),  in eastern Austria (Hundsheimer Berg) also reared from Chamaecytisus cf.  austriacus.  Egg: deposited on branch, between ridges.  Larva: very  long and slender, deeply yellow, hardly visible in mine.  Mine :  a long almost straight gallery, appearing black, at first going downwards,  often passing under the ridges and changing direction;  the mine later  fills the space between two ridges completely.  The larva is difficult or  not to be seen in the mine.  A plant with many mines can often be  recognized from a distance by the black stained branches.  Cocoon:  ochreous brown.  

Distribution: 

Widespread in Denmark.  In Sweden local in the south  (Sk. and Vg.). Not in Finland and Norway.  - Widespread in Ireland, Britain,  France, Netherlands, Belgium, West and East Germany, West and southern  Poland and Switzerland.  Also known from Spain, Portugal, Teneriffe,  Italy and Austria.  In southern Europe several undescribed species with  similar external features occur.

Life cycle: 

Voltinism:  univoltine with a long period of larval  feeding.  Larvae start feeding from September onwards.  Full-fed  larvae can be found from October until April or May;  in mild winters  most have left their mines by January, but frost can extend the period of  larval feeding considerably.  Adults on the wing from June until mid  September, in southern Europe sometimes later (October).  Life-history  and larva were described in detail by Sich (1917).


Citation: 

Description based on van Nieukerken and Johansson (1990)

Notes on description: 
Lectotype ♀ (designated here). POLAND: Silesia, Glogow (Gross Glogau). 8.vi (ante 1839) (Zeller), Walsingham coll., genitalia slide 22665 (BMNH).
Scratchpads developed and conceived by (alphabetical): Ed Baker, Katherine Bouton Alice Heaton Dimitris Koureas, Laurence Livermore, Dave Roberts, Simon Rycroft, Ben Scott, Vince Smith